小兴安岭阔叶红松林次生演替不同阶段空间结构特征研究

Study on Spatial Structure Characteristics of Broad-leaved Pinus koraiensis Forest at Different Succession Stages in the Xiaoxing'an Mountains

  • 摘要: 选取小兴安岭凉水自然保护区内阔叶红松林次生演替序列中的天然枫桦次生林、次生阔叶林、次生针阔混交林以及原始阔叶红松林为研究对象,采用空间结构参数(角尺度、大小比数、混交度)的一元和二元分布,分析林木的空间结构特征。结果表明:一元二元分布中,林分角尺度均呈现相对正态分布,在W=0.5时频率最高,4个阶段的平均角尺度分别为0.577、0.557、0.634、0.589,均以轻度聚集分布为主;并随着演替进行,聚集趋势上升,到原始阔叶红松林后聚集程度又下降;4个演替阶段林木之间呈现混交状态为极强混交,强度和极强度混交的比例依次为69.94%、70.08%、65.56%、63.64%,混交林木的比例随着演替序列呈现先上升后下降的趋势。从不同演替阶段的优势种空间分布来看,前5位优势树种呈集聚分布,树种混交程度都较高,但阳性先锋树种和耐阴树种林木的大小分化差异较大。可见,阔叶红松林4个演替阶段的变化说明演替前期林木的角尺度和混交度较大,后期尤其是原始阔叶红松林中聚集程度降低,随机分布比例上升,有利于群落长久稳定发展。

     

    Abstract: By analyzing the spatial structure characteristics of the four succession stages in the secondary succession of broad-leaved Pinus koraiensis forest, the process and trend of succession can be understood, and suggestions for the restoration and development of secondary forest can be given to speed up the succession process. At the same time, it can also give the idea of reasonable mixed management of plantation. In this paper, natural maple birch secondary forest, secondary broad-leaved forest, secondary coniferous and broad-leaved mixed forest and original broad-leaved red pine forest in the secondary succession sequence of broad-leaved red pine forest in Liangshui Nature Reserve were selected as the research object, and the spatial structure characteristics of forest trees were analyzed by using the univariate and binary distribution of spatial structure parameters (angular scale, size ratio, mixing degree). The results showed that: In the mono-binary distribution, the angular scale of the stand showed a relatively normal distribution, and the frequency was the highest when W=0.5. The average angular scale of the four stages was 0.577, 0.557, 0.634 and 0.589, respectively, and the distribution was mainly light aggregation.With the succession, the aggregation trend increased, and the aggregation degree decreased after the original broad-leaved pine forest. The size ratio of stand in the four succession stages was in the mean state, the proportion of dominant trees in the community fluctuated with succession. In the four succession stages, the trees showed extremely strong mixing, and the proportions of intensity and extreme intensity mixing were 69.94%, 70.08%, 65.56% and 63.64% in turn. The proportion of mixed forest increased first and then decreased with succession sequence. From the perspective of spatial distribution of dominant species in different succession stages, the top five dominant species showed a clustered distribution, and their tree species were highly mixed, but there was a big difference in tree size differentiation between positive pioneer species and shade tolerant species. The changes of the four succession stages of the broad-leaved Pinus koraiensis forest indicate that the angular scale and mixing degree of the trees are larger in the early succession period, and the aggregation degree decreases and the random distribution proportion increases in the later period, especially in the original broad-leaved Pinus koraiensis forest, which is conducive to the long-term and stable development of the community.

     

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