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不同比例长白落叶松–水曲柳混交林碳氮磷化学计量特征研究
Stoichiometric Characteristics of Carbon, Nitrogen and Phosphorus in Mixed Plantation of Larix olgensis and Fraxinus mandshurica under Different Mixed Proportions
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摘要: 以4种不同比例行状混交(类型Ⅰ:5∶3;类型Ⅱ:6∶4;类型Ⅲ:2∶2;类型Ⅳ:1∶1)的长白落叶松–水曲柳混交林为研究对象,测定长白落叶松和水曲柳混交林乔木叶片、凋落物和表层土壤(0~10 cm)的C、N、P元素含量,分析人工林生态系统各组分养分分配差异及循环规律。结果表明:混交比例对叶片、凋落物和土壤化学计量特征产生显著影响;叶片组分中,类型Ⅱ的N含量显著高于类型Ⅲ、Ⅳ,C∶P、N∶P显著高于其他林分类型。类型Ⅲ的P含量、C∶N显著高于类型Ⅱ;凋落物组分中,类型Ⅰ、Ⅳ的C含量、C∶N显著高于类型Ⅱ,且类型Ⅲ的C含量、C∶P、N∶P为最低。土壤组分中,类型Ⅱ具有更高的C、N、P含量和更高的C∶P、N∶P(P < 0.05);土壤养分含量与计量比对混交比例变化的响应具有趋同性,N、P含量为Ⅱ > Ⅰ > Ⅲ > Ⅳ;C∶P、N∶P为Ⅱ > Ⅲ > Ⅳ > Ⅰ。从各混交比例植物养分限制状况看,除类型Ⅱ外,其余3种林分的生长均受N元素限制。叶片、凋落物N与土壤养分含量存在强耦合关系且土壤的C、N、P含量间呈极显著正相关。综合植物生长速率、凋落物分解速率及土壤养分供应与固持能力考虑,建议在未来混交林的营造中优先选择行状比例为6∶4的水落混交林种植。Abstract: In this study, four types of Larix olgensis and Fraxinus mandshurica mixed plantations were selected as the research object according to rows-mixing ratios(type Ⅰ: 5∶3; type Ⅱ: 6∶4; type Ⅲ: 2∶2, type Ⅳ: 1∶1). We measured the C, N and P in tree leaves, litter and surface soil(0~10 cm) of L. olgensis and F. mandshurica mixed plantations to analyze the difference of nutrient allocation and cycle rule of each component in the plantation ecosystem. The results indicated that the mixed proportion had significant effects on the stoichiometric characteristics of leaves, litter and soil. In the leaf component, N content of type Ⅱ was significantly higher than that of type Ⅲ and Ⅳ, while C∶P and N∶P were significantly higher than that of the other types. P content and C∶N of type Ⅲ were significantly higher than type Ⅱ. In the litter component, C content and C∶N of type Ⅰ and Ⅳ were significantly higher than type Ⅱ, while C content, C∶P and N∶P of type Ⅲ were the lowest among all the types. In the soil component, type Ⅱ had higher C, N, P and higher C∶P, N∶P compared with the other mixed plantations(P < 0.05). The response of soil nutrient contents and stoichiometric ratios to the change of mixed proportion was convergent, N and P were Ⅱ > Ⅰ > Ⅲ > Ⅳ; C∶P and N∶P were Ⅱ > Ⅲ > Ⅳ > Ⅰ. According to the situation of plant nutrient restriction in each mixed proportion, the plantation growth was limited by N element with the exception of type II. There were strong coupling relationships between leaf, litter N and soil nutrient contents. Moreover, C, N and P in soil were significantly positively correlated. Considering the growth rate of plants, decomposition rate of litter as well as soil nutrient supply and retention capacity, our results suggested that L. olgensis and F. mandshurica plantation with row ratio of 6:4 would be preferred in the future construction of plantation.